Front-Brain – The gifts of effacer and impartation tend to manifest when the front-brain is more sprygan. In addition, those with a sprygan front-brain tend to be very communal in nature, having a strong sense of loyalty—these are not “deep” connections, rather strong bonds like those in animal packs. If a fellow dies, these sprygans will react in a practical manner as an animal does—to kill the perpetrator if it’s a threat or acknowledge that the fellow needed to be killed.

Impartation being the much rarer of the two already rare gifts, is less studied, but consistently with this we see root tendrils that exist even without head injuries, at all times. They originate in the front brain but spread into the other regions of the brain. These hybrids have a much more emotive reaction to threats against the community than other hybrids. They develop the same pack-like connections, but also develop a fiery passion for each member of their “flock,” even such members that they have only just met and know nothing of. We call this “Arbor Sickness,” as when they learn of the Nameless’ treatment, they become entirely useless to us, even when subjected to great pain. It is similar to the state of sprygans described in the histories when arbors have been burned or violated.

Low-Brain – The sensory gifts, keenears and keenscent are found when the low-brain is more sprygan. These are the most common gifts, and explains why so many hybrids have instinctual behaviors. They exhibit possessive, territorial and hunting tendencies not present in others. This makes them excellent assassins but causes some issues when we attempt to establish laws in their populous. The idea of governance or punishment of crimes is foreign to them. They adhere to “natural law.”

Upper-Brain – The two gifts of sight, heartseer and nightseer, are found when the upper-brain is more sprygan-like. In addition, for those with nightseer, it seems to provide a good sense of direction, and a distinct sense of wanting to return “home,” almost like migratory birds.

For those we have deemed Waiters—having the gifts of heartfire and impartation—when they also have nightseer, are able to guide us to the fallen stars that the Auldtree so wishes us to study. However, the effects these fallen stars have on Waiters is lethal, so great care must be taken.

A sprygan high-brain can also be associated with difficulties learning human language and an averseness to mechanics and complex tools. This can be overcome to an extent if the hybrid is properly motivated, making it imperative to force the bond between Waiters and Watchers, lest we have no leverage to bring them to their potential.

If little enough of this upper-brain region is left human, deep bonds do become difficult to cultivate. This symptom was difficult to see for some time as they are keenly aware of the difference and perfectly able to act as if they have such bonds until it no longer suits them. These bonds can still be forced, usually through isolation.

Hind-Brain – The gifts of softstep and quickstep are associated with the hind-brain being sprygan-like. A small portion of the upper hind-brain is also associated with the gifts of sight. This portion of the brain seems to allow for better coordination and allows most hybrids to “immerse” themselves in memories.